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Mark studied for a BA and MSci in Natural Sciences at Selwyn College, Cambridge, graduating in 2009. He completed a PhD in the Department of Biochemistry in Cambridge under the supervision of Dr. Daniel Nietlispach on the topic of nuclear magnetic resonance spectroscopy (NMR) method development for application to large biomolecular systems focussing particularly on membrane proteins. His PhD covered a range of topics including studying in vitro mimics of the cell membrane; protein motions by NMR; the light cycle of the photoactivated protein sensory rhodopsin II and new data processing methods for NMR spectroscopy. The latter enables experiment times to be significantly reduced. He now works as a postdoctoral researcher with Dr. Nietlispach researching protein dynamics, the role of hydrogen bonding in protein stabilisation and writing software for NMR data processing. He has taught IA Chemistry at Selwyn since 2009, and teaches IA Chemistry and IB Physical Chemistry (Chemistry A) for Churchill. He is also a keen musician and has sung with Selwyn choir for a number of years.